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Fang not a canine on Agriochoerus latifrons

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Agriochoerus latifrons (Fig 1) were recently revised when I noticed the upper fang articulated in front of the lower fang, the opposite of all other mammals. That is because that upper fang was an incisor and the upper canine was absent, creating a diastema, as in deer, elephants and other ungulates.

In addition,  the jugal (cyan) has an overlapping lacrimal (tan, Fig 1) that was previously overlooked. A similar Bauplan can be seen in Meniscotherium (Fig 2).

Figure 1. Agriochoerus latifrons with a misplaced upper fang. It’s not a canine. It’s an incisor. Also notice the lacrimal overlapping the jugal, as in Meniscotherium.

This is what happens
when you question everything. If you do so frequently enough, eventually you will learn something that others had glossed over.

Figure 2. Meniscotherium skull. In this is a smaller predecessor to Astrapotherium note the genesis of maxillary tusks here and then longer dentary when the teeth are matched to occlude correctly. Note the overlapping lacrimal.

Agriochoerus latifrons
(Leidy 1869; Late Eocene to Oligocene, 38–16mya, skeleton) has a distinctly different tooth pattern with large lateral incisors (convergent with Elephas) and a diastema. Here the canine is absent and this taxon nests with Meniscotherium.

References
Leidy J 1869. Extinct Mammals of Dakota and Nebraska: Including an account of some allied forms from other localities, together with a synopsis of the mammalian remains of North America., Journal of the Academy of Natural Sciences, Philadelphia 2(7): 135–141.

wiki/Meniscotherium
wiki/Agriochoerus


Source: https://pterosaurheresies.wordpress.com/2024/06/29/fang-not-a-canine-on-agriochoerus-latifrons/


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