Extinct Anagale compared to extant Macroscelides
When Anagale was first described
by Simpson 1931, he was faced with an enigma. He wrote, “It is not closely related to any previously known fossil insectivores,”
Figure 1. Left: Anagale skull in 3 views. Right: Macroscelides skull in 3 views.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/08/anagale-macroscelides.skull588.jpg?w=234″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/08/anagale-macroscelides.skull588.jpg?w=584″ tabindex=”0″ role=”button” class=”size-full wp-image-88468″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/08/anagale-macroscelides.skull588.jpg” alt=”Figure 1. Left: Anagale skull in 3 views. Right: Macroscelides skull in 3 views. ” width=”584″ height=”749″ />
Figure 1. Left: Anagale skull in 3 views. Right: Macroscelides skull in 3 views.
Simpson 1931 wrote:
“As regards Anagale itself, it is seen that it is much closer to the Tupaiidae than to the Macroscelididae and that it is in no sense intermediate between the two.”
Figure 2. Macroscelides proboscideus, the elephant shrew or sengis is NOT more closely related to elephants with the purported ‘Afrotheria.’ but instead is related to Tupaia, the tree shrew.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/07/macroscelides-proboscideus-recon588.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/07/macroscelides-proboscideus-recon588.jpg?w=584″ tabindex=”0″ role=”button” class=”size-full wp-image-23499″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/07/macroscelides-proboscideus-recon588.jpg” alt=”Figure 2. Macroscelides proboscideus, the elephant shrew or sengis is NOT more closely related to elephants with the purported ‘Afrotheria.’ but instead is related to Tupaia, the tree shrew.” width=”584″ height=”243″ />
Figure 2. Macroscelides proboscideus, the elephant shrew or sengis is NOT more closely related to elephants with the purported ‘Afrotheria.’ but instead is related to Tupaia, the tree shrew.
Simpson 1931 concluded,
“1. Anagale gobiensis, new genus and species, is described on the basis of remarkably well preserved skull and jaws and much of the skeleton.
2. It is shown to be closely related to the Tupaiidae (Fig 2) but referable to an extinct and non-ancestral family Anagalidae.
3. It is not closely related to any previously known fossil insectivores, none of which appears to be certainly referable to the Tupaioidea.
4. It is not intermediate between the Tupaiidae and Erinaceidae, being definitely allied with the former, but tends in some degree to strengthen the evidence of their remote common origin.
5. It tends more definitely to link the Tupaioidea and the Lemuroidea.
6. But it proves that the Tupaioidea have been distinct from all other groups of mammals from a time long antecedent to the basal Oligocene.”
In the LRT
Anagale (Fig 1) nests with an elephant shrew, Macroscelides (Fig 1) at the base of the clade that produced moles, sengis, tenrecs and odontocetes. That’s a wide gamut of morphologies, none of them arboreal. None of them close to Tupaia (Fig 2), a tree shrew.
References
Shaw G 1800. General zoology or systematic natural history. Vol. I. Part 2. Mammalia. London: Printed for G. Kearsley.
Simpson GG 1931. A new insectivore from the Oligocene, Ulan Gochu horizon, of Mongolia. American Museum Novitates 505:1-22.
wiki/Anagale
wiki/Macroscelides
Source: https://pterosaurheresies.wordpress.com/2024/09/01/extinct-anagale-compared-to-extant-macroscelides/
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