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Castoroides, the giant beaver, enters the LRT

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With the interrelationship
of beavers (Castor) and manatees recently recovered by the large reptile tree (LRT, 2330 taxa) it seemed natural to add Castoroides, the giant beaver (Figs 1–3, Foster 1838), to the matrix.

Actually the giant beaver is only about twice as long as the largest living beavers (Fig 3).
That’s big, but small compared to the recently recovered beaver descendants.

Figure 1. Skeletons of Castor, Castoroides and Moeritherium, not to scale. ” data-image-caption=”

Figure 1. Skeletons of Castor, Castoroides and Moeritherium, not to scale.

” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg?w=273″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg?w=584″ class=”size-full wp-image-91065″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg” alt=”Figure 1. Skeletons of Castor, Castoroides and Moeritherium, not to scale. ” width=”584″ height=”643″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg?w=584&h=643 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg?w=136&h=150 136w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg?w=273&h=300 273w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides_skeleton-1.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />

Figure 1. Skeletons of Castor, Castoroides and Moeritherium, not to scale.

According to Wikipedia – Castoroides
“The hind feet of the giant beaver were much larger than in modern beavers, while the hind legs were shorter.” They could grow as large as 2.2 m (7.2 ft).

Shorter legs, as in Moeritherium.
Shorter tail, as in Moeritherium.
No chevrons, as in Moeritherium.
Reduced zygomatic arch, as in Moeritherium.

And then, there’s one more thing…

Figure 1. Castoroides skull compared to scale with Castor and Moeritherium. ” data-image-caption=”

Figure 1. Castoroides skull compared to scale with Castor and Moeritherium.

” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg?w=160″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg?w=545″ class=”size-full wp-image-91054″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg” alt=”Figure 1. Castoroides skull compared to scale with Castor and Moeritherium. ” width=”584″ height=”1096″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg?w=584&h=1096 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg?w=80&h=150 80w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg?w=160&h=300 160w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/castoroides.skull588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />

Figure 1. Castoroides skull compared to scale with Castor and Moeritherium.  Note the jugal shape.

Plint, Longstaffe and Zazula 1019 reported,
“We confirm that Castoroides consumed a diet of predominantly submerged aquatic macrophytes.” = water plants.

This happens to be the diet of manatees and dugongs = big sea beavers.

In the LRT
Castoroides nests between Castor and Moeritherium, ancestral to manatees and walking sea cows like Prorastomus (Fig 3).

Figure 1. Sea cows (manatees) and their LRT ancestors to scale, presented as skeletons. ” data-image-caption=”

Figure 1. Sea cows (manatees) and their LRT ancestors to scale, presented as skeletons.

” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg?w=198″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg?w=584″ class=”size-full wp-image-91066″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg” alt=”Figure 1. Sea cows (manatees) and their LRT ancestors to scale, presented as skeletons.” width=”584″ height=”885″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg?w=584&h=885 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg?w=99&h=150 99w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg?w=198&h=300 198w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/01/sirenian_tail_evolution588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />

Figure 1. Sea cows (manatees) and their LRT ancestors to scale, presented as skeletons.

Castoroides ohioensis
(Foster 1838, Pleistocene, up to 2m long) is the giant beaver of North America. The shorter fused sacrals, tail without chevrons, shorter legs and more gracile zygomatic arch = jugal, as in Moeritherium.

The larger swimming tail of sea cows developed either 1. later as a reversal, as fossils currently indicate (see Prorastomus, Fig 3) or 2. in parallel directly from Castor (for which fossils are currently waiting to be discovered).

References
Cahn A 1932. Records and distribution of the fossil beaver, Castoroides ohioensis. J. Mammal. 12, 229–241.
Foster JW 1838. Report of Mr. Foster. In W. W. Mather (ed.), Second annual report on the geological survey of the state of Ohio
Plint T, Longstaffe FJ and Zazula G 2019. Giant beaver palaeoecology inferred from stable isotopes. Sci Rep 9, 7179 (2019). https://doi.org/10.1038/s41598-019-43710-9

mediarelations.uwo.ca/2019/05/09/giant-beavers-didnt-eat-wood-and-thats-likely-why-they-didnt-survive-the-last-ice-age/

wiki/Castoroides


Source: https://pterosaurheresies.wordpress.com/2025/01/12/castoroides-the-giant-beaver-enters-the-lrt/


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