A koala bear cousin in the Eocene: tiny Apatemys
Recent housekeeping
in the large reptile tree (LRT, 2338 taxa) brought two marsupial taxa closer together, Apatemys, the extinct apatemyid, and Phascolarctos, the much larger extant marsupial koala bear (Figs 1, 2).
Figure 1. Several views of the koala (Phascolarctos) skull to scale with that of Apatemys (see figure 2 for a larger view).
” data-image-caption=”
Figure 1. Several views of the koala (Phascolarctos) skull to scale with that of Apatemys (see figure 2 for a larger view).
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg?w=584″ class=”size-full wp-image-92819″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg” alt=”Figure 1. Several views of the koala (Phascolarctos) skull to scale with that of Apatemys (see figure 2 for a larger view).” width=”584″ height=”535″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg?w=584&h=535 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg?w=150&h=138 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg?w=300&h=275 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/koala-skull-lateral588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 1. Several views of the koala (Phascolarctos) skull to scale with that of Apatemys (see figure 2 for a larger view).All the elements in these two taxa are quite similar,
allowing for proportions to change along with the size change.
Notably Apatemys had a long tail that is lost in Phascolarctos.
Notably the orbit is very high on the skull, close to the dorsal margin in both taxa.
Notably the tooth patterns are similar.
Figure 1. The skull of Apatemys in situ and reconstructed.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg?w=584″ class=”size-full wp-image-92822″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg” alt=”Figure 1. The skull of Apatemys in situ and reconstructed. ” width=”584″ height=”559″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg?w=584&h=559 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg?w=150&h=144 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg?w=300&h=287 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/apatemys_skull_insitu588-1.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 2. The skull of Apatemys in situ and reconstructed.
These two taxa nest with arboreal
Thylacoleo and Wakaleo,which no longer nest with Phalanger in the LRT. Terrestrial wombats are widely acknowledged to be close to arboreal koalas and that is also reflected in the LRT.
Figure 2. Skeleton of the koala (genus: Phascolarctos)
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg?w=584″ class=”size-full wp-image-29981″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg” alt=”Figure 2. Skeleton of the koala (genus: Phascolarctos)” width=”584″ height=”404″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg?w=584&h=404 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg?w=150&h=104 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg?w=300&h=208 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2018/03/koala-phascolarctos-cinereus-588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 3. Skeleton of the koala (genus: Phascolarctos)
This interrelationship has been traditionally overlooked
and for good reason. Nevertheless, when taxa are tested together, as in the LRT, sometime they nest together.
Figure 1. Apatemys, only complete fossil skeleton of an apatemyid, turns out to be a basal shrew. So this clade is not extinct.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg?w=104″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg?w=356″ class=”size-full wp-image-24320″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg” alt=”Figure 1. Apatemys, only complete fossil skeleton of an apatemyid, turns out to be a basal shrew. So this clade is not extinct.” width=”584″ height=”1678″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg?w=584&h=1678 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg?w=52&h=150 52w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg?w=104&h=300 104w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2016/09/apatemys-insitu588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 4. Apatemys, only complete fossil skeleton of an apatemyid, turns out to be a basal shrew. So this clade is not extinct.
The odd manus of koalas
with two ‘thumbs’ (like a chameleon, Fig 5) is not quite reflected in the manus of Apatemys, which is odd in its own way (Fig 5) and MIGHT have been used similarly… or not.
Figure 4. The odd manus of Apatemys compared to the odder manus of the koala, Phascolarctos, which has two ‘thumbs’.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg?w=584″ class=”size-full wp-image-92925″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg” alt=”Figure 4. The odd manus of Apatemys compared to the odder manus of the koala, Phascolarctos, which has two ‘thumbs’. ” width=”584″ height=”453″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg?w=584&h=453 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg?w=150&h=116 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg?w=300&h=233 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2025/05/phascolarctos_manus_compared588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 5. The odd manus of Apatemys compared to the odder manus of the koala, Phascolarctos, which has two ‘thumbs’.
This appears to be a novel hypothesis of interrelationships.
If not, please provide a citation so I can promote it here.
References
de Blainville H 1816. Prodrome d’une nouvelle distribution systématique du règne animal. Bulletin de la Société Philomáthique, Paris (in French). 8: 113–124.
Marsh OC 1872. Preliminary description of new Tertiary Mammals. Parts I– IV: American Journal of Science 4: 122–128, 202–224.
wiki/Phascolarctos
/wiki/Koala
wiki/Apatemys
Source: https://pterosaurheresies.wordpress.com/2025/05/20/a-koala-bear-cousin-in-the-eocene-tiny-apatemys/
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