Middle Triassic Plagiosternum enters the LRT
Shoch et al 2025 wrote:
“Plagiosaurids form a small but highly disparate clade of Triassic temnospondyls that are characterized by extremely flattened and wide skulls, large orbits and a knobby to pustular ornamentation. The largest European taxon is Plagiosternum granulosum from the Middle Triassic of Germany. Originally known from fragmentary bonebed material only, recent finds add well-preserved specimens that reveal the structure of the skull, mandible and pectoral girdle in great detail. Plagiosternum granulosum was the most salt-tolerant temnospondyl in the Ladinian palaeoenvironments, where it dwelled brackish lagoons and lakes and formed a community with nothosaurids and other euhaline taxa.”
When added to
the large reptile tree (LRT, 2340 taxa) Plagiosterum nested, as expected, between two other plagiosaurs: Gerrothorax and Plagiosuchus.
Click links to see images.
The authors
“raised the question of what caused the convergent increase of orbit size in plagiosaurids.”
“The loss of circumorbital elements, which occurred convergently in up to three plagiosaurid clades (Witzmann & Schoch, 2024), might well have been triggered by different factors in divergent taxa.”
The LRT tests fewer plagiosaurids, so only one loss of circumorbital elements is indicated as the orbit increases in size, creating a confluent upper temporal fenestra by way of erosion of the postorbital elements.
“Plagiosternines, in contrast, did not have enlarged adductor muscles, as evidenced by the normal size of the adductor fenestra in the mandible.”
“Steyer (2014) suggested that salinity tolerance may have been facilitated by salt or lacrimal glands in euhaline temnospondyls, and the enormous size of the orbits in P. granulosum might well have housed such an organ.”
In the LRT
plagiosaurs are among the most primitive tetrapods, not derived temnospondyls. Add taxa to test this. Let us know if you confirm, refute or modify this LRT hypothesis.
References
Shoch RR et al (5 co-authors) 2025. Morphology and ontogeny of the plagiosaurid temnospondyl Plagiosternum granulosum from the Middle Triassic of Germany. PalZ https://doi.org/10.1007/s12542-025-00748-7
wiki/Plagiosternum
wiki/Plagiosauridae
Source: https://pterosaurheresies.wordpress.com/2026/01/31/middle-triassic-plagiosternum-enters-the-lrt/
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