Scyllacerta enters the LRT as a tiny early member of the Diapsida (within Archosauromorpha)
Scyllacerta creanae
(Jenkins et al 2026, SAM-PK-K7710a, Late Permian, South Africa) was originally classified as the oldest known member of the stem-reptile family Younginidae and the oldest neodiapsid. Adding taxa in the LRT modifies these preliminary findings. The authors did not realize a diapsid skull architecture evolved more than once by convergence and that the clade Reptilia extended back to the last common ancestor, Silvanerpeton, from the Early Cretaceous (Viséan) in the large reptile tree (LRT, 2340 taxa).
Thus the claim by the authors that ‘the origin of the tympanic fossa in reptiles is revealed by’ Scyllacerta creanae cannot be true.
Akkedops bremneri
(Mooney, Scott and Reisz, SAM-PK-K6205, Late Permian, South Africa) described as a new early reptile from South Africa. The authors included the SAM-PK-K77710a specimen in this genus. They claimed that the cranial anatomy of all three specimens was indistinguishable. The taxa are similar enough to nest together in the LRT, but the straight rostral margin of Scyllacerta is distinct from the convex rostral margin of Akkedops bremneri. Likewise these authors did not realize a diapsid skull architecture evolved more than once by convergence and that the clade Reptilia extended back to Silvanerpeton.
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References
Jenkins XA et al (10 co-authors) 2026. The origin of the tympanic fossa in reptiles revealed by a late Permian neodiapsid. Palaeontology. 69 (1) e70041: 1–15.
Mooney ED, Scott D and Reisz RR 2025. A new stem saurian reptile from the late Permian of South Africa and insights into saurian evolution. Swiss Journal of Palaeontology. 144 (1): 10.
Source: https://pterosaurheresies.wordpress.com/2026/01/30/scyllacerta-enters-the-lrt-as-a-tiny-early-member-of-the-diapsida-within-archosauromorpha/
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